The following test case produces IndexError: indices are out-of-bounds with negative integers, while positive work fine:
testdf = pd.DataFrame({"rna": (1.5,2.2,3.2,4.5), -1000: [11,21,36,40],0: [10,22,43,34], 1000:[0, 10, 20, 30]})
testdf[[1000]] #this works
testdf[[-1000]] #this does not
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