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Alcobaça Formation - Wikipedia

From Wikipedia, the free encyclopedia

Geological formation in Portugal

The Alcobaça Formation, previously known as the Guimarota Formation and also known as the Consolação Unit,[1] is a geological formation in Portugal.[2] It dates back to the Kimmeridgian stage of the Late Jurassic. It is an important source of information on the diversity of Late Jurassic mammals.[3][4] Many of the fossils were collected from the now disused and flooded Camadas de Guimarota coal mine.

Vertebrate paleofauna[edit]

Dinosaur eggs are geographically located in Lisbon District, Portugal.[5] Dinosaur tracks are geographically located in Leiria District, Portugal.[6]

Osteichthyes of the Alcobaça Formation Genus Species Location Stratigraphic Position Material Notes Images Macromesodon[7] Indeterminate Guimarota Isolated teeth Proscinetes[7] Indeterminate Guimarota Isolated teeth and prearticulars "Lepidotes"[7] Indeterminate Guimarota isolated lateral teeth, lower jaws, ganoid scales and isolated skull bones Two species of different size can be distinguished from the fossil material[7] Ionoscopidae[7] Indeterminate Guimarota Isolated teeth Macrosemiidae[7] Indeterminate Guimarota Isolated grasping tooth Caturus?[7] Indeterminate Guimarota Isolated Teeth and a partial skull Pachycormidae[7] Indeterminate Guimarota

Albanerpetontids are one of the most numerous faunal components of the Guimarota mine with around 9000 isolated remains, including parts of the skull and limbs. Remains of indeterminate salamanders and frogs are also present but far rarer.[7]

Choristoderes of the Alcobaça Formation Genus Species Location Stratigraphic position Material Notes Images Cteniogenys Indeterminate Guimarota

Indeterminate euornithopod remains located in Lisbon District.[5] Indeterminate stegosaurid remains present in Lisbon District.[5]

Ornithischians of the Alcobaça Formation Genus Species District Stratigraphic position Abundance Notes Images Dacentrurus[5] D. armatus[5] Lisboa[5] Eousdryosaurus[8] E. nanohallucis Lisboa[5] Stegosaurus[9] S. cf. ungulatus "...one tooth, five cervical vertebrae (including the axis), five dorsal vertebrae, cervical and dorsal ribs, three caudal vertebrae, chevrons, preacetabular process of the left ilium, right tibia, fibula, astragalus and [calcaneus], a cervical plate, and several fragments of plates." Trimucrodon[5] T. cuneatus[5] Lisboa[5] Later determined to be indeterminate ornithischian remains.[5]

Indeterminate sauropod remains located in Leiria and Lisboa.[10]

Mammals of the Alcobaça Formation Genus Species Location Stratigraphic position Material Notes Images Bathmochoffatia B. hapax Guimarota A paulchoffatiid multituberculate Bernardodon B. sp, B. atlanticus Porto Pinheiro A pinheirodontid multituberculate Drescheratherium D. acutum Guimarota A paurodontid dryolestidan Dryolestes D. leirensis Guimarota A dryolestid dryolestidan Ecprepaulax E. anomala Porto Pinheiro A pinheirodontid multituberculate Guimarotodon G. leiriensis Guimarota A paulchoffatiid multituberculate Guimarotodus G. inflatus Guimarota A dryolestid dryolestidan Haldanodon H. exspectatus Guimarota A docodont Henkelodon H. naias, H. sp Guimarota A paulchoffatiid multituberculate Henkelotherium H. guimarotae Guimarota A dryolestidan Iberodon I. quadrituberculatus Porto Pinheiro A pinheirodontid multituberculate Kielanodon K. hopsoni Guimarota A paulchoffatiid multituberculate Krebsotherium K. lusitanicum Guimarota A dryolestid dryolestidan Kuehneodon K. guimarotensis, K. dietrichi, K. simpsoni, K. uniradiculatus, K. sp Guimarota A paulchoffatiid multituberculate Laolestes L. andresi Porto Pinheiro A dryolestid dryolestidan Meketibolodon M. robustus Guimarota A paulchoffatiid multituberculate Meketichoffatia M. krausei, M. sp Guimarota A paulchoffatiid multituberculate Nanolestes N. krusati Porto Pinheiro A cladotherian N. drescherae Guimarota Paulchoffatia P. delgadoi Guimarota A paulchoffatiid multituberculate Pinheirodon P. sp, P. vastus, P. pygmaeus Porto Pinheiro A pinheirodontid multituberculate Plesiochoffatia P. thoas, P. peperethos Guimarota A paulchoffatiid multituberculate Portopinheirodon P. asymmetricus Porto Pinheiro A dryolestid dryolestidan Priacodon Indeterminate Porto Pinheiro A triconodontid Proalbionbaatar P. plagiocyrtus Guimarota A multituberculate of uncertain placement Pseudobolodon P. dryas, P krebsi, P. oreas Guimarota A paulchoffatiid multituberculate Renatodon R. amalthea Guimarota A paulchoffatiid multituberculate ? Tinodon Indeterminate Porto Pinheiro A tinodontid symmetrodont Xenachoffatia X. oinopion Guimarota A paulchoffatiid multituberculate Paulchoffatiidae Indeterminate Guimarota 4 distinct genera with 7 species Pinheirodontidae Indeterminate Porto Pinheiro Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited. Invertebrate Paleofauna[edit] Gastropods of the Alcobaça Formation Genus Species Location Stratigraphic Position Material Notes Images Teinostoma[7] Indeterminate Guimarota Shells Boring traces of clionidid sponges on the shells of this taxon indicate a marine lifestyle[7] Cryptaulax[7] Indeterminate Guimarota Shells Melampoides[7] M. jurassicus Guimarota Shells Boring traces of clionidid sponges on the shells of this taxon indicate a marine lifestyle, modern relatives like Melampus also venture onto land, which can also be expected from M. jurassicus[7]
  1. ^ Mateus, O.; Dinis, J.; Cunha, P. P. (2017-09-28). "The Lourinhã Formation: the Upper Jurassic to lower most Cretaceous of the Lusitanian Basin, Portugal – landscapes where dinosaurs walked". Ciências da Terra / Earth Sciences Journal. 19 (1): 75–97. doi:10.21695/cterra/esj.v19i1.355. hdl:10316/79879. ISSN 2183-4431.
  2. ^ Fürsich, Franz T.; Schneider, Simon; Werner, Winfried; Lopez-Mir, Berta; Pierce, Colm S. (2021-09-13). "Life at the continental–marine interface: palaeoenvironments and biota of the Alcobaça Formation (Late Jurassic, Central Portugal), with a formal definition of the unit appended". Palaeobiodiversity and Palaeoenvironments. 102 (2): 265–329. doi:10.1007/s12549-021-00496-x. ISSN 1867-1608.
  3. ^ "Guimarota". Paleobiology Database. Retrieved 31 August 2018.
  4. ^ "Porto Pinheiro, Lourinha". Paleobiology Database. Retrieved 31 August 2018.
  5. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
  6. ^ a b c "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
  7. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah Palmer, Douglas (September 2001). "M ARTIN , T. & K REBS , B. (eds) 2000. Guimarota. A Jurassic Ecosystem . 155 pp. München: Verlag Dr. Friedrich Pfeil. Price DM 120.00 (hard covers). ISBN 3 931516 80 6". Geological Magazine. 138 (5): 619–630. doi:10.1017/S0016756801316084. ISSN 0016-7568.
  8. ^ Fernando Escaso; Francisco Ortega; Pedro Dantas; Elisabete Malafaia; Bruno Silva; José M. Gasulla; Pedro Mocho; Iván Narváez; José L. Sanz (2014). "A new dryosaurid ornithopod (Dinosauria, Ornithischia) from the Late Jurassic of Portugal". Journal of Vertebrate Paleontology. 34 (5): 1102–1112. Bibcode:2014JVPal..34.1102E. doi:10.1080/02724634.2014.849715. S2CID 86780835.
  9. ^ Escaso, Fernando; Ortega, Francisco; Dantas, Pedro; Malafaia, Elisabete; Pimentel, Nuno L.; Pereda-Suberbiola, Xabier; Sanz, José Luis; Kullberg, José Carlos; Kullberg, María Carla (2006-12-23). "New evidence of shared dinosaur across Upper Jurassic Proto-North Atlantic: Stegosaurus from Portugal". Naturwissenschaften. 94 (5): 367–374. Bibcode:2007NW.....94..367E. doi:10.1007/s00114-006-0209-8. PMID 17187254. S2CID 10930309.
  10. ^ "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," and "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Pages 548-549.
  11. ^ Burigo, André; Mateus, Octávio (January 2025). "Allosaurus europaeus (Theropoda: Allosauroidea) Revisited and Taxonomy of the Genus". Diversity. 17 (1): 29. doi:10.3390/d17010029. ISSN 1424-2818.
  12. ^ Burigo, André; Mateus, Octávio (January 2025). "Allosaurus europaeus (Theropoda: Allosauroidea) Revisited and Taxonomy of the Genus". Diversity. 17 (1): 29. doi:10.3390/d17010029. ISSN 1424-2818.
  13. ^ "Table 13.1," in Weishampel, et al. (2004). Page 265.
  14. ^ Guillaume, A. R. D.; Puértolas-Pascual, E.; Moreno-Azanza, M. (2025). "Revisiting the choristodere and stem-lepidosaur specimens of the Guimarota Beds (Kimmeridgian, Portugal): taxonomic implications". Acta Palaeontologica Polonica. 70 (1): 77–96. doi:10.4202/app.01202.2024.
  15. ^ Malafaia, E.; Ortega, F.; Escaso, F.; Dantas, P.; Pimentel, N.; Gasulla, J. M.; Ribeiro, B.; Barriga, F.; Sanz, J. L. (2010). "Vertebrate fauna at the Allosaurus fossil-site of Andrés (Upper Jurassic), Pombal, Portugal". Journal of Iberian Geology. 36 (2): 193–204. doi:10.5209/rev_JIGE.2010.v36.n2.7. ISSN 1698-6180.

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